Eiseniella tetraedra

　ツリミミズ科 ＞ Eiseniella 属 ＞ 和名なし Eiseniella tetraedra (Savigny, 1826)

【亜種】

　　Eiseniella tetraedra tetraedra (Savigny, 1826)

　　Eiseniella tetraedra cerni Blakemore, 2004

Allurus tetraedrus Rosa, 1889: 5, 1893a: 2, 1897: 5; Michaelsen, 1890g: 305; Beddard, 1891: 269 & 270 &274, 1895: 696.

Allurus tetraëdrus Eisen, 1873: 54; Michaelsen, 1890: 12, 1891c: 207; Bretscher, 1899: 413.

Eiseniella tetraedra Michaelsen, 1900: 471, 1902a: 3 & 11; Cognetti, 1905a: 2, 1905c: 58; Gates, 1942a: 101, 1953c: 524, 1966: 244, 1967c: 160, 1976c: 2; Lindroth, 1957: 124; Heitor, 1969: 76; Martin, 1977: 308; Sims, 1978: 671; Abbott, 1982: 19; Kasprzak, 1989: 39; Schwert, 1990: 352, 354; Baha, 1997: 252; Fauci & Bezdicek, 2002: 259; Csuzdi & Zicsi, 2003: 153; Shen et al., 2005: 16; Pižl, 2006: 79; Huang et al., 2006: 18; Tsai et al., 2009: 37; Reynolds, 2010c: 132, 2011a: 259, 2015a: 116, 2015b: 181; Talavera & Pérez, 2011: 352; Szederjesi & Csuzdi, 2012: 269; Szederjesi et al., 2013a: 398, 2013b: 206; Lehmitz et al., 2014: 232; Höser, 2015: 135; Szederjesi, 2015: 147.

Allurus tetraëdra Bretscher, 1900: 39.

Eiseniella tetraedra typica Michaelsen, 1900: 472; Bretscher, 1901: 214; Cognetti, 1901b: 17, 1901a: 2, 1904c: 2, 1904g: 2, 1905e: 105, 1906a: 2, 1906d: 3, 1908d: 113, 1908e: 319; Černosvitov, 1934: 71, 1937a: 107, 1937b: 78; Tétry, 1937: 142; Easton, 1942: 245; Davies, 1954: 8; Zicsi, 1965b: 256, 1965c: 160, 1965d: 269, 1968a: 426, 1968b: 138, 1970: 241.

Helodrilus tetraedrus typica Michaelsen, 1913f: 551; Cognetti, 1913: 2; Heimburger, 1914: 284; Smith, 1917: 161, 1927: 352; Pickford, 1932: 288; Olson, 1936: 106, 1940: 2.

Helodrilus tetraedrus typicus Cognetti, 1914c: 3.

Helodrilus tetraedrus Smith, 1915: 556.

Eiseniella tetraedra tetraedra Jamieson, 1967: 70; Zicsi, 1981: 173; Easton, 1983: 481; Zicsi & Csuzdi, 1986: 120; Zicsi, 1994: 70; 2004: 126; Paliwal & Julka, 2004: 1973; Mischis & Herrera, 2006: 287; Misirlioğlu, 2008a: 470; Blakemore, 2008d: 31; Blakemore et al., 2012: 300; Stoyanović et al., 2013: 638.

Allolobophora ninnii Rosa, 1886: 1.

Allurus ninnii Rosa, 1893a: 2, 1893b: 11.

Eiseniella tetraedra ninnii Michaelsen, 1900: 472; Easton, 1942: 245.

Helodrilus tetraedrus ninnii Smith, 1917: 162.

Allurus tetragonurus Rosa, 1893a: 2.

Eiseniella tetraedra tetragonura Michaelsen, 1900: 473.

Allurus hyrcynius Rosa, 1893a: 2; Bretscher, 1899: 413.

Eiseniella tetraedra hercynia Michaelsen, 1900: 472; Černosvitov, 1937b: 79; Easton, 1942: 245; Zicsi, 1965b: 256, 1965c: 160, 1965d: 269, 1968b: 140, 1994: 71.

Helodrilus tetraedrus hercynius Smith, 1915: 556, 1917: 162, 1927: 352; Olson, 1936: 107, 1940: 2; Zicsi, 1968a: 426.

Allurus neapolitanus Rosa, 1893a: 2; Beddard, 1895: 697; Bretscher, 1899: 414.

Eiseniella tetraedra neapolitana Michaelsen, 1900: 472; Easton, 1942: 245; Easton, 1983: 481.

Helodrilus tetraedrus neapolitana Smith, 1917: 162.

Eiseniella neapolitana Szederjesi et al., 2013a: 398, 2013b: 206.

Tetragonurus pupa Beddard, 1895: 697.

Eiseniella tetraedra pupa Michaelsen, 1900: 473; Easton, 1983: 481.

Eiseniella tetraedra bernensis Michaelsen, 1900: 472; Cognetti, 1905e: 105; Zicsi, 1968a: 426.

Allolobophora tetraedra sewelli Stephenson, 1924b: 363.

Eiseniella tetraedra intermedia Zicsi, 1965b: 257, 1965c: 160, 1994: 71, 2004: 126; Easton, 1983: 481.

　　 Eiseniella tetraedra hercynia intermedia Zicsi, 1968b: 140.

Eisenia hammoniensis Easton, 1942: 245.

Eiseniella tetraedra popi Zicsi, 1968b: 140.

Eiseniella tetraedra cerni Misirlioğlu, 2008a: 469.

＜外部形態＞

　全長 30-70 mm、体幅 2-4 mm、体節数 70-100 (Zicsi, 1965c)。

　剛毛対は極めて近接し、aa:ab:bc:cd:dd = 4:1:3:1:6 (Zicsi, 1965c)。

　第一背孔は第 4/5 体節間溝 (Zicsi, 1965c)。

　かつて雄性孔と環帯の位置で forma もしくは亜種分化が認められていたが、いくつかの変種が同所的に出現することから個体変異だと考えられている (Easton, 1942; Csuzdi & Zicsi, 2003)。雄性孔は 1 つで第 11-15 体節の奇数体節に開口する。環帯は第 20、21-24、25 体節もしくは第 22、23-26、27、28、29 体節 (Easton, 1983)。思春期隆起は第 23、23 1/2-25、26 体節 (Zicsi, 1965c)。

＜内部形態＞

　隔膜は第 7/8-11/12 体節間でやや厚い (Zicsi, 1965c)。

　砂嚢は第 17 体節 (Zicsi, 1965c)。

　貯精嚢は 4 対で、第 9-12 体節 (Zicsi, 1965c)。受精囊は 2 対で、第 10-11 体節 (Zicsi, 1965c)。受精囊孔は第 9/10/11 体節間溝の cd 剛毛線と背面正中線間に開口する (Zicsi, 1965c)。

　日本国内での分布はよく分かっていないが、以下の 2 例が文献で報告されている。なお、中村 (1998) は、「アイセニア・テトラデラが北海道ウトナイ湖から」採集されたと報告しているが、形態などの情報は何も述べていないものの本種であろうと考えられる。ただし、中村 (2015) では本種の存在を認めていない。また、Blakemore et al. (2006b) は、日本初記録として富山県で採集したことを報告している。後者は形態学的特徴も含めた報告であるため確実であり、前者を参考記録として、後者を日本初記録とみなすべきであろう (南谷, 私信)。

　国外では、ポルトガル (Rosa, 1889; Heitor, 1969)、スペイン (Rosa, 1889; Zicsi, 1968a)、フランス (Cognetti, 1904g; Zicsi, 1968a)、デンマーク (Michaelsen, 1902a)、ドイツ (Michaelsen, 1890g; Lehmitz et al., 2014; Höser, 2015)、スイス (Bretscher, 1899, 1900, 1901; Zicsi, 1968a, 1970)、イタリア (Rosa, 1886; Cognetti, 1904a, 1905a, 1905e, 1914c; Zicsi, 1968a, 1981)、サルディーニャ島 (Cognetti, 1901b)、チェコ (Pižl, 2006)、オーストリア (Rosa, 1897; Zicsi, 1965b, 1965c, 1994)、ポーランド (Kasprzak, 1989)、ハンガリー (Zicsi, 1968b; Csuzdi & Zicsi, 2003)、セルビア (Stoyanović et al., 2013)、モンテネグロ (Cognetti, 1906d)、アルバニア (Szederjesi & Csuzdi, 2012)、マケドニア (Černosvitov, 1934)、ブルガリア (Černosvitov, 1937b; Zicsi & Csuzdi, 1986)、ギリシャ (Cognetti, 1913; Szederjesi, 2015)、クレタ (Cognetti, 1906d)、スウェーデン (Eisen, 1873; Michaelsen, 1902a)、ノルウェー (Eisen, 1873; Michaelsen, 1902a)、ロシア (Baturina et al., 2014)、トルコ (Misirlioğlu, 2008a)、シリア (Rosa, 1893b)、レバノン (Rosa, 1893b)、イスラエル (Rosa, 1893b; Stephenson, 1924b; Szederjesi et al., 2013b)、ヨルダン (Szederjesi et al., 2013a)、インド (Černosvitov, 1937a: 107; Paliwal & Julka, 2004)、台湾 (Shen et al., 2005; Huang et al., 2006; Tsai et al., 2009)、韓国 (Blakemore et al., 2012)、オーストラリア (Abbott, 1982)、ニュージーランド (Martin, 1977)、カナダ (Gates, 1942a; Lindroth, 1957)、アメリカ合衆国 (Heimburger, 1914; Smith, 1915, 1917, 1927; Olson, 1936, 1940; Easton, 1942; Gates, 1942a, 1953c, 1966, 1967c; Davies, 1954; Lindroth, 1957; Schwert, 1990; Fauci & Bezdicek, 2002; Reynolds, 2010c, 2011a, 2015a, b)、チリ (Michaelsen, 1890; Cognetti, 1901c, 1905c; Zicsi, 2004)、アルゼンチン (Mischis & Herrera, 2006)、トリスタン・ダ・クーニャ (Pickford, 1932)、アゾレス諸島 (Michaelsen, 1891c)、カナリア諸島 (Cognetti, 1906a; Talavera & Pérez, 2011)、カーポベルデ (Cognetti, 1908d)、アルジェリア (Baha, 1997)、南アフリカ (Cognetti, 1908e; Michaelsen, 1913f; Jamieson, 1967b; Sims, 1978) に分布する。

　半水棲種で、河岸や湖岸 (Smith, 1917; Bouche, 1972; Csuzdi & Zicsi, 2003)、小川 (Cszudi & Zicsi, 2003)、泉 (Csuzdi & Zicsi, 2003)、池の水に浸かった環境 (Smith, 1917) など、湿った環境に生息する (Easton, 1942; Csuzdi & Zicsi, 2003; Latif et al., 2009)。

　アメリカ合衆国北部では、いくつかのビニールハウス内から本種が採集された (Gates, 1966)。

　表層性種 (Lehmitz et al., 2014)。

　かつて、環帯と受精囊孔の位置から、いくつかの変種に分けられていたが、いくつもの変種が同所的に出現することから、亜種など地理的下位分類群ではなく、単なる個体変異だと考えられている (Easton, 1942; Csuzdi & Zicsi, 2003)。近年では3亜種 (tetraedra, intermedia, hercynia) が認められることが多い。また、Easton (1983) は tetraedra、intermedia、neapolitana、pupa の 4 亜種に分けている。なお、Easton (1942) によるとそれらの変種は以下の通り。なお、intermedia はホモニムであり Blakemore (2004) によって置換名 cerni が提案されている。

　　環帯 22、23-26、27 体節、雄性孔第 13 体節・・・・・・・・・・typica

　　環帯 23、23-27、28、29 体節、雄性孔第 15 体節・・・・・・・・hercynia

　　環帯 20、21-24、25 体節、雄性孔第 11 体節・・・・・・・・・・hammoniensis

　　環帯 20、21-24、25 体節、雄性孔第 13 体節・・・・・・・・・・neapolitana

　　環帯 20、21-24、25 体節、雄性孔第 15 体節・・・・・・・・・・ninnii

Allurus tetraëdrus Eisen, 1873: 54. [同定形質のみ]

Allolobophora ninnii Rosa, 1886: 1.

Allurus tetraedrus Rosa, 1889: 5.

Allurus tetraëdrus Michaelsen, 1890: 12.

Allurus tetraedrus Michaelsen, 1890g: 305.

Allurus tetraedrus Beddard, 1891: 269, 270, 274.

Allurus tetraëdrus Michaelsen, 1891c: 207.

Allurus tetraedrus Rosa, 1893a: 2 (syn. agilis, dubius).

Allurus hyrcynius Rosa, 1893a: 2.

Allurus neapolitanus Rosa, 1893a: 2.

Allurus ninnii Rosa, 1893a: 2.

Allurus tetraedrus Rosa, 1893b: 10.

Allurus ninnii Rosa, 1893b: 11.

Allurus tetraedrus Beddard, 1895: 696 (syn. agilis, novaehollandiae, dubius, hercynius).

Allurus neopolitanus Beddard, 1895: 697 (syn. ninnii).

Tetragonurus pupa Beddard, 1895: 697 (syn. tetragonurus).

Allurus tetraedrus Rosa, 1897: 5.

Allurus tetraëdrus Bretscher, 1899: 413.

Allurus hercynius Bretscher, 1899: 413.

Allurus neapolitanus Bretscher, 1899: 414.

Eiseniella tetraedra Michaelsen, 1900: 471.

Eiseniella tetraedra typica (syn. agilis, tetraedra luteus, tetraedra obscurus, amphisbaena, novis) Michaelsen, 1900: 472.

Eiseniella tetraedra hercynia Michaelsen, 1900: 472.

Eiseniella tetraedra neapolitana Michaelsen, 1900: 472.

Eiseniella tetraedra ninnii Michaelsen, 1900: 472.

Eiseniella tetraedra bernensis Michaelsen, 1900: 472 (syn. infinitesimalis).

Eiseniella tetraedra pupa Michaelsen, 1900: 473.

Eiseniella tetraedra tetragonura Michaelsen, 1900: 473.

Allurus tetraëdra Bretscher, 1900: 39.

Eiseniella tetraëdra var. typica Bretscher, 1901: 214.

Eiseniella tetraedra subsp. typica Cognetti, 1901b: 17.

Eiseniella tetraedra (typica) Cognetti, 1901c: 2.

Eiseniella tetraedra Michaelsen, 1902a: 3+11 (syn. var. lutens, var. obscurus).

Eiseniella tetraedra typica Cognetti, 1904a: 2.

Eiseniella tetraedra typica Cognetti, 1904g: 2. [記述のみ] (Oct 30, 1904)

Eiseniella tetraedra Cognetti, 1905a: 2. [記述のみ] (May 2, 1905)

Eiseniella tetraedra Cognetti, 1905c: 58.

Eiseniella tetraedra f. typica Cognetti, 1905e: 105.

Eiseniella tetraedra f. bernensis Cognetti, 1905e: 105.

Eiseniella tetraedra typica Cognetti, 1906a: 2. [記述のみ] (Feb 9, 1906)

Eiseniella tetraedra typica Cognetti, 1906d: 3. [記述のみ] (May 25, 1906)

Eiseniella tetraedra typica Cognetti, 1908d: 113. [記述のみ] (Nov 25, 1908)

Eiseniella tetraedra (typica) Cognetti, 1908e: 319. [記述のみ]

Helodrilus (Eiseniella) tetraedrus f. typica Michaelsen, 1913f: 551.

Helodrilus (Eiseniella) tetraedrus typicus Cognetti, 1913: 2. [記述のみ] (Sep 25, 1913)

Helodrilus tetraedrus var. typicus Heimburger, 1914: 284.

Helodrilus (Eiseniella) tetraedrus typicus Cognetti, 1914c: 3. [記述のみ] (Nov 3, 1914)

Helodrilus tetraedrus Smith, 1915: 556.

Helodrilus tetraedrus hercynius Smith, 1915: 556.

Helodrilus (Eiseniella) tetraedrus f. typica Smith, 1917: 161.

Helodrilus (Eiseniella) tetraedrus f. hercynia Smith, 1917: 162.

Helodrilus (Eiseniella) tetraedrus f. neapolitana Smith, 1917: 162.

Helodrilus (Eiseniella) tetraedrus f. ninnii Smith, 1917: 162.

Allolobophora (Eiseniella) tetraedra var. sewelli Stephenson, 1924b: 363.

Helodrilus tetraedrus typica Smith, 1927: 352.

Helodrilus tetraedrus hereynius Smith, 1927: 352.

Eiseniella tetraedra f. typica Pickford, 1932: 288.

Eiseniella tetraedra f. typica Černosvitov, 1934: 71.

Helodrilus (Eiseniella) tetraedrus f. typica Olson, 1936: 106.

Helodrilus (Eiseniella) tetraedrus f. hercynia Olson, 1936: 107.

Eiseniella tetraedra f. typica Tétry, 1937: 142.

Eiseniella tetraedra f. typica Černosvitov, 1937a: 107.

Eiseniella tetraedra f. typica Černosvitov, 1937b: 78.

Eiseniella tetraedra mut. hercynia Černosvitov, 1937b: 79.

Helodrilus (Eiseniella) tetraedrus f. typica Olson, 1940: 2.

Helodrilus (Eiseniella) tetraedrus f. hercynia Olson, 1940: 2.

Eiseniella tetraedra typica Easton, 1942: 245. [同定形質のみ]

Eiseniella tetraedra hercynia Easton, 1942: 245. [同定形質のみ]

Eiseniella tetraedra hammoniensis Easton, 1942: 245. [同定形質のみ]

Eiseniella tetraedra neapolitana Easton, 1942: 245. [同定形質のみ]

Eiseniella tetraedra ninnii Easton, 1942: 245. [同定形質のみ]

Eiseniella tetraedra Gates, 1942a: 101.

Eiseniella tetraedra Gates, 1953c: 524.

Eiseniella tetraedra f. typica Davies, 1954: 8.

Eiseniella tetraedra Lindroth, 1957: 124. [記述のみ]

Eiseniella tetraedra f. typica Zicsi, 1965b: 256. [記述のみ] (?, 1965)

Eiseniella tetraedra v. hercynia Zicsi, 1965b: 256. [記述のみ] (?, 1965)

Eiseniella tetraedra v. intermedia Zicsi, 1965b: 257. [記述のみ] (?, 1965)

Eiseniella tetraedra f. typica Zicsi, 1965c: 160. [形態記載] (?, 1965)

Eiseniella tetraedra v. intermedia Zicsi, 1965c: 160. [形態記載] (?, 1965)

Eiseniella tetraedra v. hercynia Zicsi, 1965c: 160. [形態記載] (?, 1965)

Eiseniella tetraedra f. typica Zicsi, 1965d: 269. [記述のみ] (?, 1965)

Eiseniella tetraedra v. hercynia Zicsi, 1965d: 269. [記述のみ] (?, 1965)

Eiseniella tetraedra Gates, 1966: 244.

Eiseniella tetraedra tetraedra Jamieson, 1967: 70.

Eiseniella tetraedra Gates, 1967c: 160.

Eiseniella tetraedra f. typica Zicsi, 1968a: 426.

Eiseniella tetraedra v. hercynia Zicsi, 1968a: 426.

Eiseniella tetraedra v. berninsis Zicsi, 1968a: 426.

Eiseniella tetraedra f. typica Zicsi, 1968b: 138. [記述のみ] (?, 1968)

Eiseniella tetraedra v. hercynia Zicsi, 1968b: 140. [記述のみ] (?, 1968)

Eiseniella tetraedra v. hercynia f. intermedia Zicsi, 1968b: 140. [記述のみ] (?, 1968)

Eiseniella tetraedra v. popi Zicsi, 1968b: 140. [記述のみ] (?, 1968)

Eiseniella tetraedra Heitor, 1969: 76. [記述のみ]

Eiseniella tetraedra f. typica Zicsi, 1970: 241.

Eiseniella tetraedra Gates, 1976c: 2. [記述のみ] (Apr, 1976)

Eiseniella tetraedra Martin, 1977: 308.

Eiseniella tetraedra Sims, 1978: 671.

Eiseniella tetraedra tetraedra Zicsi, 1981: 173. [記述のみ] (?, 1981)

Eiseniella tetraedra Abbott, 1982: 19 (syn. intermedius). [形態記載]

Eiseniella tetraedra tetraedra Easton, 1983: 481 (syn. quadrangularis, amphisbaenus, agilis, tetraedrus luteus, tetraedrus obscurus, dubius, tetragonurus syn. nov., macrurus syn. nov., flavus, tetraedrus bernensis, tetraedrus novis, tetraedrus infinitesimalis, tetraedra hammoniensis, mollis syn. nov., intermedia, tetraedra popi). [記述のみ]

Eiseniella tetraedra intermedia Easton, 1983: 481. [記述のみ]

Eiseniella tetraedra neapolitana Easton, 1983: 481 (syn. ninnii, tetraedra sewelli). [記述のみ]

Eiseniella tetraedra pupa Easton, 1983: 481 (syn. hercynius, tetraedra quadripora). [記述のみ]

Eiseniella tetraedra tetraedra Zicsi & Csuzdi, 1986: 120.

Eiseniella tetraedra Kasprzak, 1989: 39. [記述のみ]

Eiseniella tetraedra Schwert, 1990: 352, 354. [同定形質のみ]

Eiseniella tetraedra tetraedra Zicsi, 1994: 70. [記述のみ]

Eiseniella tetraedra intermedia Zicsi, 1994: 71. [記述のみ]

Eiseniella tetraedra hercynia Zicsi, 1994: 71. [記述のみ]

Eiseniella tetraedra Baha, 1997: 252.

Eiseniella tetraedra Fauci & Bezdicek, 2002: 259. [記述のみ]

Eiseniella tetraedra Csuzdi & Zicsi, 2003: 153 (syn. quadrangularis, amphisbaena, agilis, tetraedrus lutes, t. obscures, pupa, dubius, hercynius, flavus, macrurus, t. novis, t. bernensis, t. infinitesimal is, t. hammoniensis, mollis, t. popi, t. phorogenesa syn. nov., t. proporandra syn. nov.). 

Eiseniella tetraedra tetraedra Paliwal & Julka, 2004: 1973.

Eiseniella tetraedra intermedia Zicsi, 2004: 126. [記述のみ]

Eiseniella tetraedra tetraedra Zicsi, 2004: 126. [記述のみ]

Eiseniella tetraedra Shen et al., 2005: 16 (syn. hercynius).

Eiseniella tetraedra tetraedra Mischis & Herrera, 2006: 287.

Eiseniella tetraedra Pižl, 2006: 79. [記述のみ]

Eiseniella tetraedra Huang et al., 2006: 18. [記述のみ]

Eiseniella tetraedra cerni Misirlioğlu, 2008a: 469.

Eiseniella tetraedra tetraedra Misirlioğlu, 2008a: 470.

Eiseniella tetraedra tetraedra Blakemore, 2008d: 31. [同定形質のみ] (Dec ?, 2008)

Eiseniella tetraedra Tsai et al., 2009: 37.

Eiseniella tetraedra Reynolds, 2010c: 132. [記述のみ] (Nov, 2010)

Eiseniella tetraedra Talavera & Pérez, 2011: 352. [記述のみ]

Eiseniella tetraedra Reynolds, 2011a: 259. [記述のみ] (Apr, 2011)

Eiseniella tetraedra sub-spp. Blakemore, 2012b: 17. [記述のみ] (Feb 28, 2012)

Eiseniella tetraedra Szederjesi & Csuzdi, 2012: 269.

Eiseniella tetraedra tetraedra Blakemore et al., 2012: 300. [NIBR IV000021312, 243427, 213239, 213622, 213625]

Eiseniella tetraedra tetraedra Stoyanović et al., 2013: 638.

Eiseniella neapolitana Szederjesi et al., 2013a: 398.

Eiseniella tetraedra Szederjesi et al., 2013a: 398.

Eiseniella neapolitana Szederjesi et al., 2013b: 206 (syn. ninnii, tetraedra var. sewelli). [記述のみ]

Eiseniella tetraedra Szederjesi et al., 2013b: 206. [記述のみ]

Eiseniella tetraedra Lehmitz et al., 2014: 232 (syn. agilis, dubius, hercynius, tetraedra bernensis, tetraedra pupa, tetraedra tetragonura, macrura, tetraedra hammoniensis). [記述のみ]

Eiseniella tetraedra Baturina et al., 2014: 82. [記述のみ]

Eiseniella tetraedra Szederjesi, 2015: 147. [記述のみ]

Eiseniella tetraedra Höser, 2015: 135. [記述のみ]

Eiseniella tetraedra Reynolds, 2015a: 116. [記述のみ]

Eiseniella tetraedra Reynolds, 2015b: 181. [記述のみ]

[要検討]

Enterion tetraedrum Savigny, 1826: 184.

Lumbricus quadrangularis Risso, 1826: 426.

L. amphisbaena Duges, 1828: 289

Lumbricus tetraedrus Duges, 1837: 17.

Lumbricus agilis Hoffmeister 1843:191.

Lumbricus tetraedrus Eisen, 1871: 966.

Lumbricus tetraedrus luteus Eisen, 1871: 966.

Lumbricus tetraedrus obscurus Eisen, 1871: 967.

Tetragonurus pupa Eisen, 1874: 47

Allurus tetraedrus Orley, 1881: 599.

Allurus tetraedra Rosa, 1884: 51.

Allurus neapolitanus Orley, 1885: 12

Allurus tetraedrus Orley, 1885: 13.

Allolobophora ninnii Rosa, 1886: 680.

Lumbricus neapolitanus Vaillant, 1889: 113.

Lumbricus tetraedrus Vaillant, 1889: 151.

Lumbricus pupa Vaillant, 1889: 154.

Eisenia pupa Benham, 1890: 266.

Allurus tetraedrus Michaelsen, 1890: 6.

Allurus dubius Michaelsen, 1890: 6.

Allurus hercynius Michaelsen, 1890: 7.

Allurus pupa neapolitanus Michaelsen, 1890: 7.

Allurus ninnii Michaelsen, 1890: 10.

Allurus tetreaedrus Michaelsen, 1892: 251.

Allurus hercynius Michaelsen, 1892: 251. 

Allurus luteus Friend, 1892: 90

Allurus tetragonurius Friend, 1892: 242 

Allurus tetraedrus Friend, 1893: 462.

Allurus amphisbaena Friend, 1893: 462.

Allurus flavus Friend, 1893: 462.

Allurus macrurus 1893: 402.

Allurus tetragonurus Friend, 1893: 462.

Allurus tetraedrus Ribaucourt, 1896: 69.

Allolobophora tetraedrus novis Ribaucourt, 1896: 69.

Allurus tetraedrus bernensis Ribaucourt, 1896: 73. 

Allurus tetraedrus infinitesimalis Ribaucourt, 1896: 74.

Allurus tetraedrus Michaelsen, 1900: 16,19

Eiseniella macrura Michaelsen, 1900a: 474.

Eiseniella tetraedra Michaelsen, 1903: 178.

Eiseniella tetraedra hammoniensis Michaelsen, 1909: 63.

Eiseniella tetraedra var. typical Szuts, 1909: 125.

Eiseniella tetraedra var. hercynia Szuts, 1909: 126.

Allurus mollie Friend, 1911: 63.

Eisenia tetraedra intermedia Jackson, 1931: 123.

Eiseniella tetraedra typica Černosvitov, 1934: 17.

Eiseniella tetraedra intermedia Černosvitov, 1934: 17.

Eiseniella tetraedra typica Michaelsen, 1935: 40.

Eiseniella tetraedra Bodenheimer, 1935: 393.

Eiseniella tetraedra Bodenheimer, 1937: 259.

Eiseniella tetraedra ninnii Bodenheimer, 1937: 393.

Eiseniella tetraedra ninnii Cernosvitov, 1938: 549.

Eiseniella tetraedra forma? Cernosvitov, 1940: 440.

Eiseniella tetraedra quadripora Černosvitov, 1942: 220.

Eiseniella tetraedra hercynia Wilcke, 1949: 175.

Eiseniella tetraedra tetraedra Wilcke, 1949: 175.

Eiseniella tetraedra hercynia Zuck, 1951: 126.

Eiseniella tetraedra typica Zuck, 1951: 126.

Eiseniella tetraedra typica Muldal, 1952: 59.

Eiseniella tetraedra mut. hercynia Muldal, 1952: 59.

Eiseniella tetraedra Graff, 1953b: 26.

Eiseniella tetraedra f. typical Zicsi, 1959: 183.

Eiseniella tetraedra v. hercynia Zicsi, 1959: 183.

Eiseniella tetraedra popi Zicsi, 1960: 435.

Eiseniella tetraedra f. typical Zicsi, 1968: 139.

Eiseniella tetraedra f. hercynia Zicsi, 1968: 139.

Eiseniella tetraedra f. intermedia Zicsi, 1968: 140. 

Eiseniella tetraedra Gates, 1972: 108.

Eiseniella tetraedra Bouche, 1972: 215.

Eiseniella tetraedra Reynolds, 1973: 108.

Eiseniella tetraedra Reynolds, 1974: 95.

Eiseniella tetraedra Perel, 1979: 226.

Eiseniella tetraedra tetraedra Zicsi, 1982: 444.

Eiseniella tetraedra intermedia Zicsi, 1982: 444.

Eiseniella tetraedra hercynia Zicsi, 1982: 444.

Eiseniella tetraedra tetraedra Easton, 1983: 481.

Eiseniella tetraedra intermedia Easton, 1983: 480.

Eiseniella tetraedra pupa Easton, 1983: 480.

Eiseniella tetraedra Lee, 1985: 64.

Eiseniella tetraedra tetraedra Zicsi, 1991: 177.

Eiseniella tetraedra intermedia Zicsi, 1991: 178.

Eiseniella tetraedra hercynia Zicsi, 1991: 178.

Eiseniella tetraedra tetraedra Mrsic, 1991: 514.

Eiseniella tetraedra intermedia Mrsic, 1991: 514.

Eiseniella tetraedra pupa Mrsic, 1991: 514.

Eiseniella tetraedra tetraedra Qiu & Bouche, 1998: 196.

Eiseniella tetraedra pupa Qiu & Bouche, 1998: 196.

Eiseniella tetraedra intermedia Qiu & Bouche, 1998: 196.

Eiseniella tetraedra bernensis Qiu & Bouche, 1998: 196.

Eiseniella tetraedra phorogenesa Qiu & Bouche, 1998: 104.

Eiseniella tetraedra proporandra Qiu & Bouche, 1998: 105.

Eiseniella tetraedra Sims & Gerard, 1999: 90.

Eiseniella tetraedra neapolitana Pavlicek et al., 2003: 456.

Eiseniella tetraedra tetraedra Pavlicek et al., 2003: 457.

 Eiseniella tetraedra Csuzdi & Pavlicek, 2005: 91.

Eiseniella neapolitana Csuzdi & Pavlicek, 2005: 91.

Michaelsen, 1907:228, Michaelsen, 1911:142, Jackson, 1931:123, Blakemore, 2008: 41-42.

Abbott, I., 1982. The distribution of earthworms in the Perth Metropolitan area. Rec. West. Aust. Mus. 10(1): 11-34.

Baturina, M., Timm, T., Loskutova, O., 2014. Oligochaeta (Annelida, Clitellata) communities in lakes of the Ural Mountains (Russia). Zoosymposia 9: 77-94.

Baha, M., 1997. The earthworm fauna of Mitidja, Algeria. Tropical Zoology 10: 247-254.

Beddard, F.E., 1891. The classification and distribution of earthworms. Proceedings of the Royal Physical Society of Edinburgh 10: 235-290.

Beddard, F.E., 1895. A Monograph of the order of Oligochaeta. Oxford at the Clarendon Press. pp. 769.

Blakemore, R.J., 2008d. A review of Japanese earthworms after Blakemore (2003). In: Ito MT, Kaneko N, (eds.), A Series of Searchable Texts on Earthworm Biodiversity, Ecology and Systematics from Various Regions of the World, 2nd Edition (2006) and Supplemental. COE soil Ecology Research Group, Yokohama National University, Japan. CD-ROM Publication.

Blakemore, R.J., 2012b. Japanese earthworms revisited a decade on. Zoology in the Middle East 58 suppl 4: 15-22.

Blakemore, R.J., Cho, J.L., Park, T.S., 2012. Six exotic terrestrial earthworms (Oligochaeta: Megadrilacea: Moniligastridae, Lumbricidae, Ocnerodrilidae & Megascolecidae) newly added to Korean species biodiversity list. Zootaxa 3368: 300-304.

Bretscher, K., 1899. Beitrag zur Kenntnis der Oligochaeten-fauna der Schweiz. Revue Suisse de Zoologie 6: 369-426.

Bretscher, K., 1900. Mitteilungen über die Oligochaetenfauna der Schweiz. Revue Suisse de Zoologie 8: 1-44.

Bretscher, K., 1901. Beobachtungen über Oligochæten der Schweiz. Revue Suisse de Zoologie 9: 189-223.

Černosvitov, L., 1934. Die Lumbriciden Bulgariens. Mitteilungen aus den königlichen Naturwissenschaftlichen Instituten in Sofia 7: 71-78.

Černosvitov, L., 1937a. On a collection of Indian earthworms of the family Lumbricidae. Records of the Indian Museum 39(2): 105-111.

Černosvitov, L., 1937b. Die Oligochatenfauna Bulgariens. Mitteilungen aus den königlichen Naturwissenschaftlichen Instituten in Sofia 10: 69-92.

Cognetti, L., 1901b. Res Italicae. III. Gli Oligocheti della Sardegna. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 16(404): 1-26.

Cognetti, L., 1901c. Oligocheti raccolti dal Dott. F. Silvestri nel Chile e nella Repubblica Argentina. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 16(407): 1-2.

Cognetti. L., 1904a. Nota su alcuni Lombricidi di caverne italiane. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 19(459): 1-4.

Cognetti, L., 1904g. Lombricidi dei Pirenei. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 19(476): 1-14.

Cognetti, L., 1905a. Oligocheti dell'isola d'Elba e di Pianosa. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 20(490): 1-6.

Cognetti, L., 1905c. Gli Oligocheti della regione Neotropicale. Mem. R. Acad. Soc. Torino 56: 1-72.

Cognetti, L., 1905e. Lombrichi liguri del Museo Civico di Genova. Annali del Museo Civico di storia naturale di Genova 42: 102-127.

Cognetti, L., 1906a. Contributo alla conoscenza della drilofauna delle isole Canarie. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 21(521): 1-4.

Cognetti, L., 1906d. Nuovi dati sui Lumbricidi dell'Europa orientale. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 21(527): 1-18.

Cognetti, L., 1908d. Lombrichi raccolti dal cav. Leonardo Fea nelle isole del Capo Verde e nel golfo di Guinea. Annali del Museo Civico di storia naturale di Genova 4: 79-118.

Cognetti, L., 1908e. Di alcuni Oligocheti esotici. Annalen des Naturhistorischen Museums in Wien 22(4): 308-321.

Cognetti, L., 1913. Escursioni zoologiche del Dr. E. Festa nell Isola di Rodi. V. Oligocheti dell'Isola di Rodi. Bollettino del Laboratorio di zoologia generale e agraria della R. Scuola superiore d'agricultura in Portici 28(674): 1-6.

Cognetti, L., 1914c. Escursioni Zoologiche del Dr. Enrico Festa nei monti della Vallata del Sangro (Abruzzi). Nota sugli Oligocheti degli Abruzzi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 29(689): 1-5.

Csuzdi, C., Zicsi, A., 2003. Earthworms of Hungary (Annelida: Oligochaeta; Lumbricidae). Pedozoologica Hungarica 1 1: 1-112.

Davies, H., 1954. A preliminary list of the earthworms of northern New Jersey with notes. Breviora, Museum of Comparative Zoology 26: 1-13.

Easton, E.G., 1983. A guide to the valid names of Lumbricidae (Oligochaeta). In: Satchell JE (ed.), Earthworm Ecology: from Darwin to Vermiculture, Chapman and Hall, London, pp. 475-487.

Easton, T.H. jr., 1942. Earthworms of the northeastern United States: a key, with distribution records. Journal of Washington Academy of Sciences 32(8): 242-249.

Eisen, G., 1873. Om Skandinaviens Lumbricider. Ofversigt af Kongl. Vetenskaps-Akademiens Förhandlingar 30 (8): 43-56.

Fauci, M.F., Bezdicek, D.F., 2002. Lumbricid earthworms in the Palouse region. Northwest Science 76(3): 257-260.

Gates, G.E., 1942a. Check list and bibliography of north American earthworms. American Midland Naturalist 27(1): 86-108.

Gates, G.E., 1953c. On the earthworms of the Arnold Arboretum, Boston. Bulletin of the Museum of Comparative Zoology at Harvard College 107(10): 500-534.

Gates, G.E., 1966. Requiem~ for megadrile utopias. A contribution toward the understanding of the earthworm fauna of North America. Proceedings of the Biological Society of Washington 79: 239-254.

Gates, G.E., 1967c. On the earthworm fauna of the great American desert and adjacent areas. Great Basin Nat. 27: 142-176.

Gates, G.E., 1976c. More on oligochaete distribution in North America. Megadrilogica 2(11): 1-6.

Heimburger, H.V., 1914. Notes on Indiana earthworms. Proceedings of the Indiana Academic Sciences 1914: 281-285.

Heitor, P.F.P.C., 1969. Les Lombricidés du Portugal. Pedobiologia 9: 76-80.

Höser, N., 2015. Zur Regenwurmfauna (Oligochaeta: Lumbricidae), insbesondere zur Proctodrilus- Verteilung, in den Auen der Tschernosem- und Parabraunerde- Gebiete beiderseits des Erzgebirges. Opuscula Zoologica, Budapest 46(2): 133-142.

Huang, J., Qin, X., Zheng, J.S., Chong, W., Dong, M.Z., 2006. Research on earthworm resources of China: I. Check-list and distribution. Journal of China Agricultural University 11(3): 9-20.

Jamieson, B.G.M., 1967b. New records of Ocnerodrilidae and Lumbricidae (Oligochaeta) from South African rivers. Zoologica Africana 3: 59-86.

Kasprzak, K., 1989. Zoogeography and habitat distribution of earthworms (Lumbricidae) and Enchytraeids (Enchytraeidae) of the Carpathian mountains (Poland). Miscellania Zoologica 13: 37-44.

Lehmitz, R., Römbke, J., Jänsch, S., Krück, S., Beylich, A., Graefe, U., 2014. Checklist of earthworms (Oligochaeta: Lumbricidae) from Germany. Zootaxa 3866(2): 221-245.

Lindroth, C.H., 1957. The Faunal Connections between Europe and North America. John Wiley & Sons, Inc., NY. 344 pp.

Martin, N.A., 1977. Guide to the lumbricid earthworms of New Zealand pastures. New Zealand Journal of Experimental Agriculture 5: 301-309.

Michaelsen, W., 1890. Oligochaeten des Naturhistorischen Museums in Hamburg. Mitteilungen aus dem Naturhistorischen Museum in Hamburg 6/7: 2-13.

Michaelsen W., 1890g. Oligochaeten des Naturhistorischen Museums in Hamburg. IV. Jahrbuch der Hamburgischen Wissenschaftillchen Anstalten 8: 299-340.

Michaelsen, W., 1900. Oligochaeta. Das Tierrich 10: 1-575. [in German]

Michaelsen, W., 1902a. Die Lumbriciden-Fauna Norwegens und ihre Beziehungen. Abdruck Verhandelingen der Naturwissenschaftlichen Vereins zu Hamburg 9(3): 1-13.

Michaelsen, W., 1913f. Die Oligochäten des Kaplandes. Zoologische Jahrbücher, Abteilung für Systematik, Jena 34: 473-556.

de Mischis, C.C., Herrera, J.A.D., 2006. Review of the distribution of exotic earthworms (Annelida, Oligochaeta) in Argentina and confirmed examples of their introduction. Carribean Journal of Science 42(3): 285-293.

Misirlioğlu, M., 2008a. Some earthworm records from Anatolia (Oligochaeta, Lumbricidae). Turkish Journal of Zoology 32: 469-471.

中村好男, 1998. ミミズと土と有機農業. 創森社, 123 p.

Olson, H.W., 1936. Earthworms of Missouri. Ohio Journal of Science 36(2): 102-113.

Olson, H.W., 1940. Earthworms of New York State. American Museum Novitates 1090: 1-9.

Paliwal, R., Julka, J.M., 2004. Checklist of earthworms of western Himalaya, India. Zoos' Print Journal 20(9): 1972-1976.

Pižl V, 2006. Earthworms (Oligochaeta, Lumbricidae) of Kokořínsko Protected Landscape Area. Bohemia Centralis, Praha 28: 75-83. [in Polishi ?]

Reynolds, J.W., 2010c. The earthworms (Oligochaeta: Acanthodrilidae, Lumbricidae, Megascolecidae and Sparganophilidae) of northeastern United States, revisited. Megadrilogica 14(7): 101-157.

Reynolds, J.W., 2011a. The earthworms (Oligochaeta: Acanthodrilidae, Eudrilidae, Glossoscolecidae, Komarekionidae, Lumbricidae, Lutodrilidae, Megascolecidae, Ocnerodrilidae, Octochaetidae and Sparganophilidae) of southeastern United States. Megadrilogica 14(9-12): 175-318.

Reynolds, J.W., 2015a. A checklist by counties of earthworms (Oligochaeta: Lumbricidae, Megascolecidae and Sparganophilidae) in the states of Connecticust, Massachusetts and Rhode Island, USA. Megadrilogica 17(8): 113-123.

Reynolds, J.W., 2015b. A checklist by counties of earthworms (Oligochaeta: Lumbricidae, Megascolecidae and Sparganophilidae) in New Jersey, USA. Megadrilogica 17(12): 179-190.

Rosa, D., 1886. Note sui lombrici del Veneto. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 1(3): 1.

Rosa, D., 1889. Note sui Lombrichi iberici. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 4(63): 1-5.

Rosa, D., 1893a. Catalogo e distribuzione geografica dei Lumbricidi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 8(151): 1-5.

Rosa, D., 1893b. Viaggio del Dr. E. Festa in Palestina, nel Libano e regioni vicine. II. Lumbricidi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 8(160): 1-14.

Rosa, D., 1897. Nuovi Lombrichi dell'Europa orientale (Seconda serie). Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Università di Torino 12(269): 1-5.

Schwert, D.P., 1990. Oligochaeta: Lumbricidae. In: Dindal DL (ed.), Soil Biology Guide, John Wiley & Sons, NY. pp. 341-356.

Shen, H.P., Tsai, S.C., Tsai, C.F., 2005. Occurence of the earthworms Pontodrilus litoralis (Grube, 1855), Metaphire houlleti (Perrier, 1872) and Eiseniella tetraedra (Savigny, 1826) from Taiwan. Taiwania 50(1): 11-21.

Sims, R.W., 1978. Megadriacea (Oligochaeta). In: Werger MJA (ed), Biogeography and Ecology of Southern Africa. Monographiae Biologicae, No. 31. pp. 661-676.

Smith, F., 1915. Two new varieties of earthworms with a key to described species in Illinois. Bulletin of the Illinois State Laboratory of Natural History 10: 551-559.

Smith, F., 1917. North American earthworms of the family Lumbricidae in the collections of the United States National Museum. Proceedings of the United States National Museum 52: 157-182.

Smith, F., 1928. An account of changes in the earthworm fauna of illinois and a description of one new species. Bulletin of the Illinois Natural History Survey 17:347-362.

Stephenson, J., 1924b. On some Indian Oligochaeta with a description of two new genera of Ocnerodrilinae. Records of the Indian Museum 26: 317-365.

Stoyanović, M., Tsekova, R., Pešic, S., Milutinović, J., Milutinović, T., 2013. Diversity and a biogeographical review of the earthworms (Oligochaeta: Lumbricidae) of the Balkan Mountains (Stara Planina Mountains) in Serbia and Bulgaria. Turkish Journal of Zoology 37: 635-642.

Szederjesi, T., 2015. New earthworm records from various parts of Greece (Oligochaeta: Lumbricidae, Acanthodrilidae, Megascolecidae, Ocnerodrilidae). Opuscula Zoologica, Budapest 46(2): 143-152.

Szederjesi, T., Scuzdi, C.S., 2012. New earthworm species and records from Albania (Oligochaeta, Lumbricidae). Acta Zoologica Academiae Scientiarum Hungaricae 58(3): 259-274.

Szederjesi T, Pavlíček T, Csuzdi Cs, 2013a. New earthworm species from Jordan (Oligochaeta, Lumbricidae). Acta Zoologica Academiae Scientiarum Hungaricae 59(4): 391-400.

Szederjesi, T., Pavlíček, T., Csuzdi, C.S., 2013b. New data to the earthworm fauna of Israel (Oligochaeta, Lumbricidae). Opusc. Zool. Budapest 44(2): 201-209.

Talavera, J.A., Pérez, D.I., 2011. Earthworms in relict natural woodlands on the westernmost Canary Islands: species richness and ecological preferences. Systematics and Biodiversity 9(4): 349-360.

Tétry, A., 1937. Révision des lombriciens de la collection de Savigny. Bulletin Museum national d’Histoire Naturelle 9(2): 140-155.

Tsai, C.F., Shen, H.P., Tsai, S.C., Lin, K.J., Hsieh, H.L., Yo, S.P., 2009. A checklist of oligochaetes (Annelida) from Taiwan and its adjacent islands. Zootaxa 2133: 33-48.

Zicsi, A., 1965b. Beiträge zur Kenntnis der Lumbricidenfauna Österreichs. Opuscula Zoologica, Budapest 5(2): 247-265.

Zicsi, A., 1965c. Die Lumbriciden Oberösterreichs und Österreichs unter Zugrundlegung der Sammlung Karl Wesselys mit besonderer Berücksichtigung des Linzer Raumes. Naturkundliches Jahrbuch der Stadt Linz 11: 125-201.

Zicsi A, 1965d. Bearbeitung der Lumbriciden-Sammlung des Naturhistorischen Museums von Wien. Opuscula Zoologica, Budapest 5(2): 267-272.

Zicsi, A., 1968a. Revision der Regenwurm-Sammlung des Naturhistorischen Museums von Genf. Revue Suisse de Zoologie 75: 419-433.

Zicsi, A., 1968b. Ein zusammenfassendes Verbreitungsbild der Regenwürmer auf Grund der Boden- und Vegetationsverhältnisse Ungarns. Opuscula Zoologica, Budapest 8(1): 99-164.

Zicsi, A., 1970. Revision der Bretscherischen Regenwurm-Sammlung aus Zürich. Revue Suisse de Zoologie 77: 237-246.

Zicsi, A., 1981. Wietere Angaben zur Lumbricidenfauna Italiens (Oligochaeta: Lumbricidae). Opuscula Zoologica, Budapest 17/18: 157-180.

Zicsi, A., 1994. Die Regenwürmer Österreichs (Oligochaeta: Lumbricidae) mit Bestimmungstabellen der Arten. Verhandlungen der Zoologisch-Botanischen Gesellschaft in Wien 131: 37-74.

Zicsi, A., 2004. Die Regenwurmfauna Chiles mit Bestimmungsschlüsseln der Chilota-, Yagansia- und Eodrilus- Arten (Oligochaeta: Acanthodrilidae) Regenwürmer aus Südamerika 40. Berichte des naturwissenschaftlichen-medizinischen Verein Innsbruck 91: 111-128.

Zicsi, A., Csuzdi, C., 1986. Regenwürmer aus Bulgarien (Oligochaeta: Lumbricidae). Opuscula Zoologica Budapest 22: 113-121.