Aporrectodea caliginosa (Savigny, 1826)

　ツリミミズ科＞バライロツリミミズ属

Allolobophora caliginosa Rosa, 1893a: 1, 1893b: 7, 1897: 4; Michaelsen, 1894a: 182, 1897: 68, 1899b: 222; Beddard, 1895: 699, 1896: 196; Stephenson, 1924b: 363; Pickford, 1932: 288; 小林, 1941d: 376; Easton, 1942: 246; Gates, 1953a: 2, 1953c: 505, 1958a: 2; Davies, 1954: 2; Lindroth, 1957: 123; Zicsi, 1965b: 261, 1965c: 175, 1968a: 428, 1968b: 153, 1969: 75, 1970: 243, 1981: 173, 1994: 63; Heitor, 1969: 76; Martin, 1977: 303; Righi, 1984b: 119; Pop & Pop, 2006: 1221.

Helodrilus caliginosus Michaelsen, 1902a: 3&11; Ude, 1905: 497; Cognetti, 1906d: 6, 1908d: 113, 1914c: 3; Heimburger, 1914: 285.

Aporrectodea caliginosa Gates, 1942a: 102; Sims, 1978: 671; Easton, 1981: 41; Abbott, 1982: 15; Rushton & Luff, 1984: 18; Schwert, 1990: 355; Springett, 1992: 1378; Mischis & Righi, 1999: 64; Blakemore, 2003: 9, 2008d: 27, 2012b: 17; Zicsi, 2004: 124; Pižl, 2006: 78; Huang et al., 2006: 18; Hong & Kim, 2009: 137; Tsai et al., 2009: 37; Mischis & Herrera, 2006: 287; Sawar et al., 2006: 4; Chang et al., 2009: 16; Talavera & Pérez, 2011: 352; Blakemore, 2013: 39;上平, 2013: 43; Stoyanović et al., 2013: 637; Szederjesi et al., 2013a: 392, 2013b: 201; 2014b: 89; Szlavecz et al., 2014: 318; Shekhovtsov et al., 2014: 53-2; Lehmitz et al., 2014: 224; Misirlioğlu & Szederjesi, 2015: 100; Szederjesi, 2015: 144; Höser, 2015: 135.

Nicodrilus caliginosus Baha, 1997: 249.

Helodrilus caliginosus typicus [sic] Michaelsen, 1900: 483.

Helodrilus caliginosus typica Michaelsen, 1908: 245, 1913f: 551, 1922: 14; Smith, 1917: 166; Olson, 1936: 108, 1940: 4.

Allolobophora caliginosa typica Stephenson, 1923: 500, 1931: 89; Tétry, 1937: 146; Černosvitov, 1937b: 85; Kobayashi, 1940b: 295, 1941b: 150; 小林, 1941c: 264, 1941g: 515.

Aporrectodea caliginosa caliginosa Easton, 1983: 476; Zicsi & Csuzdi, 2001: 139; Paliwal & Julka, 2004: 1972; Christoffersen, 2011: 145; Szederjesi, 2014: 49; Shekhovtsov et al., 2018: 10.

Allolobophora caliginosa caliginosa Dhiman & Battish, 2006: 2135.

Nicodrilus caliginosus caliginosus Zeriri et al., 2013: 231.

Allolobophora beddardi Michaelsen, 1894a: 182.

Allolobophora turgida Eisen, 1873:46; Ude, 1894: 58; Friend, 1910: 81; Gates, 1966: 242, 1967c: 153.

Allolobophora caliginosa turgida Bretscher, 1899: 414, 1900: 39.

Helodrilus (Allolobophora) borellii Cognetti, 1904g: 2.

Allolobophora borellii Baha, 1997: 251.

Helodrilus (Helodrilus) mariensis Stephenson, 1917: 414.

Allolobophora (Eophila) mariensis Stephenson, 1923: 504.

Allolobophora arnoldi Gates, 1952a: 1, 1953a: 2, 1953c: 510; Davies, 1954: 1.

Allolobophora molita Gates, 1952a: 3, 1953c: 513.

Aporrectodea caliginosa alternisetosa Easton, 1983: 477.

Aporrectodea caliginosa meridionalis Easton, 1983: 477.

Aporrectodea caliginosa obscuricola Easton, 1983: 477.

[再チェック]

Allolobophora caliginosa trapezoides Song & Paik, 1969: 13.

＜外部形態＞

　全長 35-200 mm (35-200 [Chang et al., 2009]、60 [Cognetti, 1904g])、体幅 3.8-6 mm (3 [Cognetti, 1904g]、3.5-4 [Chang et al., 2009])、体節数 125-160 (117-246 [Chang et al., 2009]、125-160 [Kobayashi, 1941b]、132 [Cognetti, 1904g])。

　背面は白灰色、ピンク色から暗褐色 (Kobayashi, 1941b)。またはピンク色から明褐赤色 (Chang et al., 2009)。環帯は肉色、灰色または褐灰色 (Kobayashi, 1941b)、暗色 (Chang et al., 2009)。

　口前葉は上口前葉 (Chang et al., 2009)。剛毛は1体節2本4対。第一背孔は 6/7〜14/15 体節間溝。

　環帯は鞍型で、第 25-35 体節 (25-35 [Cognetti, 1904g]、26, 27-34 [Kobayashi, 1941b]、27、28、29、30-34、35 [Chang et al., 2009]) を占める。受精嚢孔は 2 対で第 9/10/11 体節間孔の cd 剛毛線に開口する (Chang et al., 2009)。雌性孔は第 14 体節に 1 対で、b 剛毛線に開口し、小さなスリット状 (Chang et al., 2009)。雄性孔は第 15 体節に 1 対で大きなスリット状で、b 剛毛線と c 剛毛線の間に開口する (Chang et al., 2009)。性的隆起壁は第 31-34 1/2 体節 (Cognetti, 1904g)、または第 31 と 33 体節 (Chang et al., 2009)。ab 剛毛周辺の剛毛隆起壁は第 9-11、27、30、32-34、35 体節にある (Chang et al., 2009)。

＜内部形態＞

　第 5/6/7/8/9/10 体節間の隔膜はやや厚い (Chang et al., 2009)。

　砂嚢は第 17 体節 (Chang et al., 2009)。石灰腺は第 10 体節に 1 対 (Chang et al., 2009)。腸管は第 20 体節から膨れる (Chang et al., 2009)。

　心臓は第 6-11 体節 (Chang et al., 2009)。

　受精嚢は 2 対で、第 10-11 体節で丸くて小さい (Chang et al., 2009)。腎管は巨大腎管 (Chang et al., 2009)。卵巣は第 13 体節 (Chang et al., 2009)。清掃は 2 対で、第 10-11 体節 (Chang et al., 2009)。貯精嚢は第 9-12 体節 (Chang et al., 2009)。摂護腺を欠く (Chang et al., 2009)。

　国外ではイギリス (Friend, 1910; Rushton & Luff, 1984)、ポルトガル (Heitor, 1969)、フランス (Cognetti, 1904g; Stephenson, 1931; Zicsi, 1968a)、オランダ (Michaelsen, 1922)、デンマーク (Michaelsen, 1902a)、ドイツ (Lehmitz et al., 2014; Höser, 2015)、スイス (Bretscher, 1899, 1900; Zicsi, 1968a)、イタリア (Cognetti, 1914c; Zicsi, 1981)、チェコ (Pižl, 2006)、オーストリア (Rosa, 1897; Zicsi, 1965b, 1969, 1994)、ポーランド (Kasprzak, 1989)、ハンガリー (Zicsi, 1968b; Csuzdi & Zicsi, 2003)、クロアチア (Cognetti, 1906d)、セルビア (Stoyanović et al., 2013)、モンテネグロ (Szederjes, 2014)、ルーマニア (Pop & Pop, 2006; Szederjesi et al., 2014b)、ブルガリア (Rosa, 1897; Černosvitov, 1937b)、ギリシャ (Szederjesi, 2015)、クレタ島 (Cognetti, 1906d)、ノルウェー (Michaelsen, 1902a)、スウェーデン (Michaelsen, 1902a)、ロシア (Shekhovtsov et al., 2014)、トルコ (Misirlioğlu & Szederjesi, 2015)、シリア (Rosa, 1893b)、レバノン (Rosa, 1893b)、イスラエル (Szederjesi et al., 2013b)、ヨルダン (Rosa, 1893b; Szederjesi et al., 2013a)、イラン (Latif et al., 2009)、パキスタン (Stephenson, 1917)、インド (Michaelsen, 1908; Stephenson, 1923, 1924b; Gates, 1958a; Paliwal & Julka, 2004; Sawar et al., 2006; Dhiman & Battish, 2006)、台湾 (Huang et al., 2006; Tsai et al., 2009; Chang et al., 2009)、中国 (Kobayashi, 1940b)、韓国 (Song & Paik, 1969; Hong & Kim, 2009; Blakemore, 2013)、オーストラリア (Abbott, 1982)、ニュージーランド (Michaelsen, 1899b; Martin, 1977; Springett, 1992)、トンガ (Ude, 1905)、ハワイ諸島 (Beddard, 1896)、カナダ (Gates, 1942a; Lindroth, 1957)、アメリカ合衆国 (Michaelsen, 1894a; Ude, 1894; Heimburger, 1914; Olson, 1936, 1940; Easton, 1942; Gates, 1942a, 1952a, 1953a, 1966, 1967c; Davies, 1954; Lindroth, 1957; Schwert, 1990; Szlavecz et al., 2014)、メキシコ (Gates, 1942a)、コスタリカ (Gates, 1942a)、ベネズエラ (Christoffersen, 2011)、コロンビア (Christoffersen, 2011)、エクアドル (Christoffersen, 2011)、ペルー (Christoffersen, 2011)、ブラジル (Christoffersen, 2011)、ボリビア (Christoffersen, 2011)、パラグアイ (Righi, 1984b)、チリ (Zicsi & Csuzdi, 2001; Zicsi, 2004; Christoffersen, 2011)、アルゼンチン (Righi, 1984b; Mischis & Herrera, 2006; Christoffersen, 2011)、トリスタン・ダ・クーニャ (Pickford, 1932)、カナリー諸島 (Talavera & Pérez, 2011)、カーポベルデ (Cognetti, 1908d)、エジプト (Michaelsen, 1897)、モロッコ (Michaelsen, 1897)、アルジェリア (Michaelsen, 1897; Baha, 1997; Zeriri et al., 2013)、チュニジア (Michaelsen, 1897)、南アフリカ (Michaelsen, 1913f; Sims, 1978) に分布する。

　北方四島（国後島、色丹島）にも分布するが、分子系統学的解析から人為的に移入されたものだと考えられている (Shekhovtsov et al., 2018)。

　土壌有機物の少ない（< 4 %）(Jänsch et al., 2013, Lehmitz et al., 2014 から二次引用)、農耕地や草原に生息する (Smith, 1917; Springett, 1992; Csuzdi & Zicsi, 2003; Lehmitz et al., 2014)。また、庭園 (Csuzdi & Zicsi, 2003)、森林 (Smith, 1917; Bösener, 1965, Lehmitz et al., 2014 から二次引用; Csuzdi & Zicsi, 2003) にも生息する。なお、砂地 (Csuzdi & Zicsi, 2003)、川底 (Smith, 1917) でも発見される。カナリー諸島では、人出のあまり入っていない天然林内にも普通に生息している (Talavera & Pérez, 2011)。ニュージーランドの農地では、最も出現頻度の高い種であった (Springett, 1992)。

　アメリカ合衆国北部のグリーンハウス内から採集記録がある (Gates, 1966)。

　Boucheによる生態区分では、地中種 endogeics とされている (Bouche, 1972; Csuzdi & Zicsi, 2003; Lehmitz et al., 2014)。しばしば、地表に大量の糞塊を排出する (Csuzdi & Zicsi, 2003)。

　撹乱環境下では、短期間で在来種と置き換わってしまうことが報告されている (Miller et al., 1955, Csuzdi & Zicsi, 2003 から二次引用)。

　かつて和名はカッショクツリミミズまたはクロイロツリミミズとされたが、＃＃＃による A. caliginosa 種群の分類学的再検討により、本種は体色を欠くものを指し、黒色もしくは褐色に着色するものは A. trapezoides を指すとされたため、これらの和名は本種の和名として不適切である。このため、新たな和名の指定が必要であり、種名不明とした。

　世界的に広く分布し、単為生殖型・多様な倍数体を含んでいるため、様々な種名で記載されている。Csuzdi & Zicsi (2003) はそれらの多数の種の再検討が必要であるとして、それまでは trapezoides や turgida、tuberculata などの細分化を認めないとしている。なお、彼ら自身はいくつかの亜種・種に細分化されるだろうと予測している。また、Szederjesi et al. (2013b) や Lehmitz et al. (2014) は trapezoides を本種のシノニムとしている。Easton (1983) は caliginosa、alternisetosa、meridionalis、obscuricola、trapezoides の 5 亜種を認めている。

　Csuzdi & Zicsi (2003) は、monticola や carochensis、tetramuralis を本種のシノニムとした。

Allolobophora turgida Eisen, 1873: 46 (syn. cyaneum, communis cyaneus). [形態記載]

Allolobophora caliginosa Rosa, 1893a: 1 (syn. trapezoides, turgida).

Allolobophora caliginosa Rosa, 1893b: 7 (syn. trapezoides).

Allolobophora caliginosa Michaelsen, 1894a: 182.

Allolobophora beddardi Michaelsen, 1894a: 182.

Allolobophora turgida Ude, 1894: 57.

Allolobophora caliginosa Beddard, 1895: 699 (syn. trapezoides, communis, turgida, novaae-hollandiae).

Allolobophora caliginosa Beddard, 1896: 196.

Allolobophora caliginosa Ribaucourt, 1896: 48 (syn. anatomicus [part], communis [part], turgida, cyanea, novaeholandiae, australiensis).

Allolobophora caliginosa Rosa, 1897: 4.

Allolobophora caliginosa Michaelsen, 1897: 68.

Allolobophora caliginosa Michaelsen, 1899b: 222.

Allolobophora caliginosa var. turgida Bretscher, 1899: 414.

Helodrilus (Allolobophora) caliginosus (typicus) Michaelsen, 1900: 483.

Allolobophora caliginosa var. turgida Bretscher, 1900: 39.

Helodrilus (Allolobophora) caliginosus Michaelsen, 1902a: 3+11.

Helodrilus (Allolobophora) borellii Cognetti, 1904g: 2. [形態記載] (Oct 30, 1904)

Helodrilus (Allolobophora) caliginosus Ude, 1905: 497.

Helodrilus (Allolobophora) caliginosus Cognetti, 1906d: 6. [記述のみ] (May 25, 1906)

Helodrilus (Allolobophora) caliginosus f. typica Michaelsen, 1908: 245.

Helodrilus (Allolobophora) caliginosus Cognetti, 1908d: 113. [記述のみ] (Nov 25, 1908)

Allolobophora turgida Friend, 1910: 81.

Helodrilus (Allolobophora) caliginosus f. typica Michaelsen, 1913f: 551.

Helodrilus caliginosus Heimburger, 1914: 285.

Helodrilus (Allolobophora) caliginosus Cognetti, 1914c: 3. [記述のみ] (Nov 5, 1914)

Helodrilus (Helodrilus) mariensis Stephenson, 1917: 414.

Helodrilus (Allolobophora) caliginosus f. typica Smith, 1917: 166.

Allolobophora (Allolobophora) caliginosa f. typica Michaelsen, 1922: 14.

Allolobophora (Allolobophora) caliginosa f. typica Stephenson, 1923: 500.

Allolobophora (Eophila) mariensis Stephenson, 1923: 504.

Allolobophora (Allolobophora) caliginosa Stephenson, 1924b: 363.

Allolobophora caliginosa (typica) Stephenson, 1931: 89.

Helodrilus (Allolobophora) caliginosus f. typica Olson, 1936: 108.

Allolobophora caliginosa f. typica Černosvitov, 1937b: 85.

Allolobophora caliginosa Pickford, 1938: 288.

Allolobophora caliginosa f. typica Tétry, 1937: 146.

Helodrilus (Allolobophora) caliginosus f. typica Olson, 1940: 4.

Allolobophora caliginosa f. typica Kobayashi, 1940b: 295. [形態記載]

Allolobophora caliginosa f. typica Kobayashi, 1941b: 150. [形態記載]

Allolobophora caliginosa typica 小林, 1941c: 264. [記述のみ]

Allolobophora caliginosa 小林, 1941d: 376. [記述のみ]

Allolobophora caliginosa typica 小林, 1941g: 515. [記述のみ]

Allolobophora caliginosa Easton, 1942: 246 (part). [同定形質のみ]

Aporrectodea caliginosa Gates, 1942a: 102.

Allolobophora arnoldi Gates, 1952a: 1.

Allolobophora molita Gates, 1952a: 3.

Allolobophora arnoldi Gates, 1953a: 2.

Allolobophora caliginosa Gates, 1953a: 2.

Allolobophora caliginosa Gates, 1953c: 505.

Allolobophora arnoldi Gates, 1953c: 510.

Allolobophora molita Gates, 1953c: 513.

Allolbophora arnoldi Davies, 1954: 1.

Allolobophora caliginosa Davies, 1954: 2.

Allolobophora caliginosa Linderoth, 1957: 123. [記述のみ]

Allolobophora caliginosa Gates, 1958a: 2.

Allolobophora caliginosa Zicsi, 1965b: 261. [記述のみ] (?, 1965)

Allolobophora caliginosa Zicsi, 1965c: 175. [形態記載] (?, 1965)

Allolobophora turgida Gates, 1966: 242.

Allolobophora turgida Gates, 1967c: 153.

Allolobophora caliginosa Zicsi, 1968a: 428.

Allolobophora caliginosa Zicsi, 1968b: 153. [記述のみ] (?, 1968)

Allolobophora caliginosa Heitor, 1969: 76. [記述のみ]

Allolobophora caliginosa trapezoides Song & Paik, 1969: 13.

Allolobophora caliginosa Zicsi, 1969: 75. [記述のみ] (?, 1969)

Allolobophora caliginosa Zicsi, 1970: 243.

Allolobophora caliginosa Martin, 1977: 303.

Aporrectodea caliginosa Sims, 1978: 671.

Aporrectodea caliginosa Easton, 1981: 39. [同定形質のみ] (Apr 30, 1981)

Allolobophora caliginosa Zicsi, 1981: 173. [記述のみ] (?, 1981)

Aporrectodea caliginosa Abbott, 1982: 15 [形態記載].

Aporrectodea caliginosa Easton, 1983: 476 (syn. carneum, lividus, gordianus, purus, hortensiae, novaehollandiae, communis olivaceus, communist pellucidus, turgida turgida, turgida tuberculata, levis, australiensis, inflata, borellii, similis, remyi, caliginosa hellenica, nocturna, iowana, arnoldi, molt). [記述のみ]

Aporrectodea alternisetosa Easton, 1983: 477. [記述のみ]

Aporrectodea meridionals Easton, 1983: 477. [記述のみ]

Aporrectodea obscuricola Easton, 1983: 477. [記述のみ]

Aporrectodea caliginosa Rushton & Luff, 1984: 18. [記述のみ]

Allolobophora caliginosa Righi, 1984b: 119. [記述のみ] (??, 1984)

Allolobophora caliginosa Kasprzak, 1989: 39. [記述のみ]

Aporrectodea caliginosa Schwert, 1990: 355 (syn. turgida). [同定形質のみ]

Aporrectodea caliginosa Springett, 1992: 1378. [記述のみ]

Allolobophora caliginosa Zicsi, 1994: 63. [記述のみ]

Nicodrilus caliginosus Baha, 1997: 249.

Allolobophora borellii Baha, 1997: 251.

Aporrectodea caliginosa Mischis & Righi, 1999: 64.

Aporrectodea caliginosa caliginosa Zicsi & Csuzdi, 2001: 139. [記述のみ]

Aporrectodea caliginosa Csuzdi & Zicsi, 2003: 75 (syn. craneum, trapezoides, gordianus, lividus, purus, capensis, hortensiae, novaehollandiae, olivaceus, pellucidus, turgida, turgida tuberculata, levis, australiensis, beddardi, inflata, borellii, similis, maeriensis, remyi, augilensis, obscuricola, caliginosa hellenica, samarigera graeca, nocturna, iowans, arnoldi, molita, caliginosus paratypicus, caliginosus meridionalis, caliginosus meridionalis pseudolongus, caliginosus alternisetosus, monticola syn. nov., carochensis syn. nov., tetramuralis syn. nov.). [形態記載]

Aporrectodea caliginosa Blakemore, 2003: 9. [記述のみ]

Aporrectodea caliginosa caliginosa Paliwal & Julka, 2004: 1972.

Aporrectodea caliginosa Zicsi, 2004: 124 (syn. caliginosa trapezoides). [記述のみ]

Aporrectodea caliginosa Pižl, 2006: 78. [記述のみ]

Aporrectodea caliginosa Hong & Kim, 2009: 137.

Aporrectodea caliginosa Tsai et al., 2009: 37.

Aporrectodea caliginosa Mischis & Herrera, 2006: 287.

Aporrectodea caliginosa Sawar et al., 2006: 4.

Allolobophora caliginosa caliginosa Dhiman & Battish, 2006: 2135.

Allolobophora caliginosa Pop & Pop, 2006: 1221.

Aporrectodea caliginosa Huang et al., 2006: 18. [記述のみ]

Aporrectodea caliginosa Blakemore, 2008d: 27. [記述のみ] (Dec ?, 2008)

Aporrectodea caliginosa Chang et al., 2009: 16. [形態記載] (2009)

Aporrectodea caliginosa Latif et al., 2009: 2 (syn. trapezoides) (part). [同定形質のみ]

Aporrectodea caliginosa caliginosa Christoffersen, 2011: 145.

Aporrectodea caliginosa Talavera & Pérez, 2011: 352. [記述のみ]

Aporrectodea caliginosa Blakemore, 2012b: 17. [記述のみ] (Feb 28, 2012)

Aporrectodea caliginosa Blakemore, 2013: 39.

Aporrectodea caliginosa 上平, 2013: 43.

Aporrectodea caliginosa Stoyanović et al., 2013: 637.

Nicodrilus caliginosuscaliginosus [sic] Zeriri et al., 2013: 231.

Aporrectodea caliginosa Szederjesi et al., 2013a: 392.

Aporrectodea caliginosa (part.) Szederjesi et al., 2013b: 201 (syn. caliginosa var trapezoides). [記述のみ]

Aporrectodea caliginosa Szlavecz et al., 2014: 318 (syn. turgida). [記述のみ] (Jan 2014)

Aporrectodea caliginosa caliginosa Szederjesi, 2014b: 49. [記述のみ] (Jun 2014)

Aporrectodea caliginosa Szederjesi et al., 2014b: 89. [記述のみ] (Jun 2014)

Aporrectodea caliginosa Shekhovtsov et al., 2014: 52-2. [記述のみ] (Aug 2014)

Aporrectodea caliginosa (part) Lehmitz et al., 2014: 224 (syn. cranium, inflate, caliginosa trapezoides, nocturnus). [記述のみ] (23 Sep 2014)

Aporrectodea caliginosa Szederjesi, 2015: 144. [記述のみ]

Aporrectodea caliginosa Höser, 2015: 135. [記述のみ]

Aporrectodea caliginosa Misirlioğlu & Szederjesi, 2015: 100 (syn. turgida). [記述のみ] (Jul 2015)

Aporrectodea caliginosa caliginosa Shekhovtsov et al., 2018: 10. [記述のみ]

[non Aporrectodea caliginosa]

Michaelsen, 1907: 229 (=trapezoides by Abbott, 1982).

Michaelsen, 1901: 142 (=trapezoides by Abbott, 1982).

Jackson, 1931: 126 (=trapezoides by Abbott, 1982).

大石, 1934: 104 (= trapezoides by Abbott, 1982).

Easton, 1942: 246, Stop-Bowitz, 1969: 191.

[要検討]

Enterion caliginosum Savigny, 1826: 180.

Enterion carneum Savigny, 1826: 180.

Lumbricus trapezoides Dugés, 1828: 289.

Lumbricus gordianus Templeton, 1836: 235.

Lumbricus lividus Templeton, 1836: 235.

Lumbricus purus Dugés, 1837: 17.

Lumbricus communis anatomicus Hoffmeister, 1845 (part): 28. (non Lumbricus anatomicus Duges, 1828 = Allolobophora chlorotica chlorotica (Savigny, 1826)).

Lumbricus hortensiae Kinberg, 1867: 98.

? Lumbricus helenae Kinberg, 1867: 98.

Lumbricus novaehollandiae Kinberg, 1867: 99.

Lumbricus capensis Kinberg, 1867: 100. 

Lumbricus olivaceus Eisen, 1871: 964.

Lumbricus pellucidus Eisen, 1871: 964.  (non L. pellucidus Templeton, 1834)

Allolobophora turgida Eisen, 1873: 46.

Allolobophora turgida tuberculata Eisen, 1874: 43.

Lumbricus levis Hutton, 1877: 351.

Allolobophora turgida Örley, 1881: 595.

Lumbricus cyaneus Vejdovsky, 1883: 228.

Lumbricus laevis Vejdovsky, 1883: 228.

Lumbricus levis (part) Vejdovsky, 1883: 228.

Lumbricus australiensis Fletscher, 1886: 539.

Allolobophora (Allolobophora) caliginosa Rosa 1893a: 424.

Allolobophora beddardi Ribaucourt, 1896: 40.

Allolobophora inflata Michaelsen, 1899: 124.

Helodrilus borellii Cognetti, 1904: 2.

Helodrilus (Allolobophora) caliginosus: Szüts, 1909: 135.

Allolobophora similis Friend, 1910: 99.

Allolobophora remyi Černosvitov, 1929: 149.

Eophila augilensis Sciacchitano, 1931: 302.

Allolobophora caliginosa typica Cordero, 1931: 353.

Allolobophora caliginosa Bodenheimer 1935: 393.

Helodrilus caliginosus Bodenheimer 1937: 259.

Eophila obscuricola Černosvitov, 1936: 5.

Dendrobaena samarigera graeca Černosvitov, 1938: 191.

Allolobophora caliginosa Pop, 1943: 13.

Allolobophora caliginosa hellenica Tzelepe, 1943: 1.

Allolobophora nocturna Evans, 1946: 98.

Allolobophora iowana Evans, 1948: 515.

Allolobophora caliginosa: Zicsi 1968: 153.

Nicodrilus (Nicodrilus) caliginosus caliginosus paratypicus Bouché, 1972: 327.

Nicodrilus (Nicodrilus) caliginosus alternisetosus Bouché, 1972: 333.

Nicodrilus (Nicodrilus) caliginosus meridionalis Bouché, 1972: 334.

Nicodrilus (Nicodrilus) caliginosus meridionalis pseudolongus Bouché, 1972: 335.

Allolobophora turgida (part) Gates, 1972a: 84.

Allolobophora tuberculata Gates, 1972a: 79.

Allolobophora australiensis Reynolds & Cook, 1976: 74.

Helodrilus caliginosum Reynolds & Cook, 1976: 84.

Allolobophora purus Reynolds & Cook, 1976: 160.

Nicodrilus caliginosus: Perel, 1979: 209.

Allolobophora caliginosa: Zicsi, 1982: 425.

Aporrectodea trapezoides Gates, 1982: 28.

Aporrectodea tuberculata Gates, 1982: 28.

Aporrectodea turgida Gates, 1982: 28.

Allolobophora caliginosa: Zicsi, 1991 185.

Aporrectodea (Aporrectodea) caliginosa: Mršić, 1991: 321.

Aporrectodea caliginosa caliginosa Perel, 1997: 63.

Aporrectodea caliginosa caliginosa Zicsi & Csuzdi, 1999: 996.

Nicodrilus (Nicodrilus) caliginosus caliginosus Qiu & Bouché, 1998: 188.

Nicodrilus (Nicodrilus) caliginosus meridionalis Qiu & Bouché, 1998: 188.

Nicodrilus (Nicodrilus) noctunus (laps.) Qiu & Bouché, 1998: 188.

Nicodrilus monticola Pérez & Rodriguez, 2002: 517.

Nicodrilus carochensis Pérez & Rodriguez, 2002: 520.

Nicodrilus tetramuralis Pérez & Rodriguez, 2002: 521.

Aporrectodea caliginosa Blakemore, 2002: 273.

Aporrectodea caliginosa caliginosa Pavlicek et al., 2003: 456.

Aporrectodea caliglinosa Csuzdi & Pavlicek, 2005: 88.

Allolobophora caliginosa Beddard, 1896: 62, Michaelsen, 1899a: 27, Righi, 1979: 146, 1984a: 119, Zicsi, 1993: 638.

Allolobophora turgida Eisen, 1874:46, Ljungstrom et al., 1973: 240.

Aporrectodea caliginosa (part) Gerard, 1964: 27, Sims & Gerard, 1985: 54.

Aporrectodea caliginosa caliginosa Easton, 1983: 476, Blakemore, 2006: 2.

Aporrectodea caliginosa trapezoides Zicsi & Csuzdi, 1988: 217, 2001: 139, Zicsi, 1993: 638.

Aporrectodea turgida Reynolds, 1977: 56, Gates, 1977: 56, Mischis, 1999: 24, Zicsi & Csuzdi, 2001: 139.

Helodrilus (Allolobophora) caliginosa Michaelsen, 1899c: 27, 1904: 288, Santelices et al., 1973: 67.

Lumbricus communis cyaneus (part) Hoffmeister, 1845: 28. (non Enterion cyaneum Savigny, 1826), Hoffmeister, 1845: 28.

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